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Tim completed his PhD in molecular toxicology at Colorado State University, where he made use of flow cytometric techniques and developed multimethod computational models to understand heterogenous cellular states in response to different xenobiotics and in degenerative aging. Tim joined the Spencer Lab in June 2019 to delve deeper into single-cell quantifications and to investigate the various mechanisms and stress responses underlying proliferative cellular heterogeneity. Tim completed his post-doc in the Spencer Lab in August 2023 to take a position as Senior Scientist for Calico Life Sciences in San Fransico, CA.